The Pyrgotidae are an unusual family of flies (Diptera), one of only two families of Cyclorrhapha that lack ocelli. Most species are "picture-winged" (i.e, have patterns of bands or spots on the wings), as is typical among the Tephritoidea, but unlike other tephritoids, they are endoparasitoids; the females pursue scarab beetles in flight, laying an egg on the beetle's back under the elytra where the beetle cannot reach it. The egg hatches and the fly larva...