Some observations of Herpeteros angelus, 1/26/25
Surveyors: @arthroverts, @lexiechambie, @fowlivia, @tokkioka, @paperplum, myself. Under the auspices of @invertclubsocal.
We met upon a hillside in the Santa Clarita area of Los Angeles County about 3:15 PM, ready to begin our survey for Soledad Desertsnails (Herpeteros angelus). We walked out upon the hilltop, and for the next hour and a half searched extensively, looking under rocks, debris and dead yucca (Hesperoyucca whipplei) on both sides of the ridge. No snails of any kind were forthcoming, though several California Trapdoor Spider (Bothriocyrtum californicum) burrows were observed.
We decided to relocate, and at approximately 5:00 PM, arrived at the base of another hillside a short distance away.
This location was at the southern end of the Sierra Pelona Mountains, where the foothills began giving way to the Santa Clarita Valley. According to Macrostrat data, the terrain was composed of “Unconsolidated clay, silt and sand colluvial deposits on hillsides, commonly containing abundant angular rock fragments…”
The surrounding habitat was defined by relatively steep, rolling slopes covered in mixed chaparral. Immediately evident plant species included California Buckwheat (Eriogonum fasciculatum), Chaparral Yucca (Hesperoyucca whipplei), and grasses. Some of this vegetation was clearly burned, evidence of past conflagrations in this fire-prone area (noted by @davegoodward in a comment on one of my observations of H. angelus). Scattered stones lay about, in highest density on top of the ridge.
It had rained less than 24 hours prior, and the ground and vegetation were damp. Clouds covered much of the sky, though there was no precipitation the whole time we were there. The ambient temperature remained in the low 40s F, measured at 44° at 5:15 PM.
At approximately 5:05 PM, I observed the first living snail of the evening, perched on a blackened yucca leaf within the rosette of a dead Hesperoyucca whipplei. The elevation on this south-facing slope was roughly 1,600 feet. Temperature and weather conditions were consistent with the initial assessment. At this time, the sun was about ten minutes from setting, though hidden behind the clouds.
The specimen was likely a subadult. Its shell measured approximately 13mm in diameter and had a dark band around the body whorl. The amber ground color of the shell was broken up by black striations on the apical surface, turning to spots in umbilical view. The lip of the shell, and the dorsum of the foot, were covered in the orange mucus that, according to a comment by @davegoodward on the previously mentioned observation, is diagnostic for Herpeteros angelus. The tentacles were gray-toned and darker, consistent with Gregg’s description of H. angelus.
After photographing this specimen, I carefully returned it to the dead yucca where I had found it. Several yards to the southwest, @arthroverts and @lexiechambie had discovered a second, much smaller snail deep within a dead yucca rosette. This juvenile’s size and the poor light conditions prevented me from obtaining detailed photographs. It had also retracted into its shell, so we were not able to observe the body. However, this snail did appear much like the first specimen, possessing a band and dark apical striations. However, it differed in its size (less than a centimeter shell diameter) and in possessing very obvious papillae all over the apical shell surface; perhaps a characteristic of very young individuals.
In the meantime, @fowlivia and @tokkioka had discovered two more snails. After photographing the other specimens, I hiked up to the prominence where the pair of adults were concealed in a small cavity under a rock, arriving at about 5:30 PM. At this elevated, sloping summit, grasses were dominant and rocks were strewn about in higher density than on the adjacent lower slopes.
At this juncture, the sun had been down for about 15 minutes and dusk had arrived. The weather and temperature conditions were still essentially identical as at first assessment.
The two snails were touching tentacles, head to head, perhaps as part of a courtship ritual. They were slightly larger than the first specimen, with shell diameters of perhaps 15-20mm (though I regrettably neglected to measure them). Their shells were more pale than that of the first snail, likely a consequence of age. However, the cream-colored apical surface gave way to pale amber with black striations, as it did on the subadult. The chestnut band was present on the body whorl, and the tentacles were darker than the pearly gray dorsal surface of the foot where it entered the aperture (though this gave soon way to an orange tone, due to the mucus, on most of the foot). The foot of the subadult, in contrast, was almost entirely orange. The shell lips were orange, as in the subadult.
Aside from these living individuals, two empty shells were observed, one of which I photographed. This larger of the two shells was found by @fowlivia under vegetation.
One other snail species was observed in the immediate vicinity of this last observation: two empty shells of Rumina decollata, an introduced, predatory Achatinid snail. This was a discouraging sign for the native snails we had just observed. Aside from the threat of human development and climate change, they now must contend with this voracious predator of other gastropods.
Sadly, the complete absence of snails from our first survey site may represent the extirpation of H. angelus from that locality.
However, the fact of this species’ continued survival, in an area relentlessly dominated and partitioned by modern human development, is hopeful if also astounding. The hillside we ascended was the terminus of one narrow finger of wild land, surrounded on three sides by houses, businesses, and roads. The contrast between the natural darkness of the hillside and the bright lights of the strip just below was blatant, and somewhat absurd. How many people in this city knew that just a short way up from where the road now passed, a unique form of life had evolved and was still holding steadfast after hundreds of thousands of years, at the least?
Development; now predatory invasives; and more and more, climate change. The Soledad Desertsnail has lasted this long, but if it is to survive, we must work to undo and slow the effects of modern human expansion in this tiny corner of the still-living world.
References
Alvarez, R.M. and K.R. Bovard, 2005, Preliminary Geologic Map of the Los Angeles 30' x 60' Quadrangle, Southern California: U.S. Geological Survey Open-File Report 2005-1019. 112 / 2665309, 2662641
(Via Macrostrat, https://macrostrat.org/)
Gregg, W. O., 1948, “A New and Unusual Helicoid Snail from Los Angeles County, California.” Bulletin of the Southern California Academy of Sciences, 47(3), 100-102.