Notes from a visit to Kosierskraal Game Farm (Agulhas Plain, Western Cape, South Africa) in 2001, part 2

...continued from https://www.inaturalist.org/journal/milewski/80491-notes-from-a-visit-to-kosierskraal-game-farm-agulhas-plain-western-cape-south-africa-in-2001#

DISCUSSION

I find scant reference to Kosierskraal on the Web today, suggesting that the game farm is now defunct, at least as a tourist-attraction.

However, it is now one of the sites for the quagga re-breeding project (https://www.quaggaproject.org/ and https://www.inaturalist.org/observations/113812038 and https://www.inaturalist.org/observations/115584188). This is confirmed by https://www.inaturalist.org/observations/101930627), despite the misspelling 'Coetzeekraal'.

Kosierskraal is, indeed, one of three locations for the quagga re-breeding project, in and near the Agulhas Plain:

Nuwejaars wetland (https://nuwejaars.com/ and https://www.6000.co.za/tag/nuwejaars-wetlands/), on the Agulhas Plain, south of Kosierskraal:

https://www.inaturalist.org/observations/101322717
https://www.inaturalist.org/observations/20161644
https://www.inaturalist.org/observations/108503112

Northwest of Napier, on the other side of Soetmuisberg (https://en.wikipedia.org/wiki/Napier,_South_Africa) :

https://www.inaturalist.org/observations/115898967

The special interest of this region, for me, was the combination of fynbos (which is regarded as unpalatable and unproductive for large animals) and the incidence of considerable populations of wild ungulates in prehistoric times on the Agulhas Plain.

It is remarkable that the ungulates at Kosierskraal showed no interest in supplementary nutrients, in the form of lick-blocks, despite resorting to osteophagy. Furthermore, melanism in the common eland seems to indicate the opposite of the expected deficiency in copper.

In July 2001, I received an email from Mick D'alton, summarising what he later told me in person:
"During times of drought and subsequent food shartage in the past, we have experienced mortalities among the eland and bontebok in particular. All investigations have concluded that death was from high internal parasite infestations, without identifying a cause for the infestation. As copper shortage was suggested as a contributing factor, I tried on many occasions - without success - to get them to take a copper lick, both in block and powder form. So, to date there has been no mineral supplementation".

I was also surprised that

  • the black wildebeest fared as well as it did, given that the Agulhas Plain is far from its recent distribution, and
  • the bontebok fared poorly, despite this being its typical habitat.

The black wildebeest at Kosierskraal foraged mainly on previously cultivated land. However, it remained true that, over a considerable period here, it was actually the black wildebeest that fared better than the truly indigenous bontebok - as tested by the lack of forage of quality during drought, and parasites.

I infer the possibility that these two alcelaphin grazers competed with each other, to the detriment of the bontebok. I suspect that, if Mick D'alton had not introduced the black wildebeest, the bontebok might have fared well at Kosierskraal.

It is clear that the common eland fares well on the coastal plains of the southwestern Cape, even when its original seasonal movements are no longer possible. Furthermore, it grows to its full genetic potential here, in contrast to populations conserved in the Drakensberg (https://hikingthedrakensberg.blogspot.com/2017/09/wildlife-of-drakensberg-story-of-eland.html and https://en.wikipedia.org/wiki/Drakensberg), where stunted-looking individuals have been typical (https://www.inaturalist.org/observations/64785188 and https://www.inaturalist.org/observations/10007389 and https://www.inaturalist.org/observations/11332783 and https://www.inaturalist.org/observations/92906834).

Another surprise was that, even in the Fynbos biome, and even south of Bredasdorp, Trinervitermes is an important ('herbivorous') termite, capable of supporting the largest specialised termitophage on Earth, the aardwolf. (Please see https://www.inaturalist.org/journal/milewski/59182-comparisons-of-termites-and-termite-eating-animals-in-africa-and-australia-part-1#.)

This seems linked to ungulates, in the sense that Trinervitermes depends on the mildness of the fire regime. Under a regime of occasional, intense wildfire, the termitaria are destroyed, and graminoids, particularly grasses, are excluded.

Posted on June 18, 2023 01:21 AM by milewski milewski

Comments

Puzzled by the success of the black wildebeest when introduced to the Fynbos Biome, I phoned Graham Avery (https://www.researchgate.net/profile/Graham-Avery-2 and https://independent.academia.edu/GrahamAvery and https://africanscientists.africa/business-directory/avery/ and https://za.linkedin.com/in/graham-avery-b3b91419) in late August 2001.

He told me the following:

There is no palaeontological evidence for the occurrence of the black wildebeest in the southwestern part of South Africa in the Holocene. Thus, the success of this species on game farms and nature reserves in Western Cape province is unexplained.

During the glacial maximum (18-20 thousand years ago), the black wildebeest, like the springbok (Antidorcas marsupialis), was probably centred on the Karoo, but may have extended on to the broadly exposed continental shelf.

At Die Kelders cave on the Agulhas Plain, the most recent records of the black wildebeest are 50-60 thousand years old. The maximum incidence seems to have been 50-90 thousand years ago.

Graham Avery suspected that this species persisted near Stanford/Gansbaai until about 30 thousand years ago, despite the lack of evidence of such occurrence on the Agulhas Plain at the last glacial maximum.

'Glacial' deposits up to about 20 thousand years ago, at Swartklip (https://www.cambridge.org/core/journals/quaternary-research/article/abs/paleoanthropological-implications-of-the-nonarcheological-bone-assemblage-from-swartklip-i-southwestern-cape-province-south-africa/27EC82614E8D1A4C4052C1AD9F306EB4 and https://za.geoview.info/swartklip,3360798) and Sea Harvest (https://journals.co.za/doi/pdf/10.10520/AJA00382353_9146 and https://www.researchgate.net/figure/Map-showing-the-location-of-MSA-Sea-Harvest-and-Hoedjiespunt-and-LSA-Lynch-Point_fig1_272568764), on the western coastal foreland, include wildebeest, reedbuck (Redunca), and other grazers. The deposits of bones in the dens of hyaenas include the teeth of a large form of jackal, indicating cold climates.

Graham Avery doubted that any warthog (Phacochoerus) occurred on this western coastal foreland at the time of European colonisation. Molars of this genus have been found at Sea Harvest, dating to 65-85 thousand years ago.

He also thought that the quagga (Equus quagga) probably did not visit the Swartland (https://upload.wikimedia.org/wikipedia/commons/f/fe/Map_of_the_Western_Cape_with_Swartland_highlighted_%282011%29.svg and https://en.wikipedia.org/wiki/Swartland) seasonally. It is not that invoking a migration of large ungulates to the western coastal area of the southwestern Cape is far-fetched; it is that there is not much evidence of the occurrence of equids in this region during the Holocene.

Posted by milewski 11 months ago

Common grasses in coastal renosterveld (https://en.wikipedia.org/wiki/Swartland_Shale_Renosterveld), in its original state, include Themeda triandra (https://www.inaturalist.org/taxa/208571-Themeda-triandra), Ehrharta calycina (https://www.inaturalist.org/taxa/76766-Ehrharta-calycina), Hyparrhenia hirta (https://www.inaturalist.org/taxa/77479-Hyparrhenia-hirta), and Cymbopogon marginatus (https://www.inaturalist.org/taxa/583194-Cymbopogon-marginatus).

In similar vegetation on Signal Hill (https://en.wikipedia.org/wiki/Signal_Hill_(Cape_Town)), the main grasses, about 30 years ago, were Themeda triandra, Hyparrhenia hirta, and Tenaxia disticha (https://www.inaturalist.org/taxa/595151-Tenaxia-disticha).

Reference:
Stock et al. (1993, https://inis.iaea.org/search/search.aspx?orig_q=RN:24060928)

Posted by milewski 10 months ago

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